Agnes Arber

She was the first woman to receive the Gold Medal of the Linnean Society of London. Her scientific research focused on the monocotyledon group of flowering plants. She also contributed to development of morphological studies in botany during the early part of the 20th century. Her later work concentrated on the topic of philosophy in botany, particularly on the nature of biological research.

• Name — Agnes Arber • Birth Name — Agnes Robertson • Born — 23 February 1879 • Birthplace — Primrose Hill, London, England • Died — 22 March 1960 • Place of death — Cambridge, England, UK • Fields — Plant morphology, plant anatomy • Education — University College London, (BSc, 1899) Newnham College (1902) University College, London (Sc.D., 1905) • Author Abbrev Bot — A.Arber • Awards — Linnean Medal of the Linnean Society of London (1948) • Spouse — Edward Alexander Newell Arber (m. 1909) • Children — Muriel Agnes (1913–2004)

Agnes Robertson was born on 23 February 1879 in Primrose Hill, London. She was the first child of Henry Robert Robertson, an artist, and Agnes Lucy Turner, and had three younger siblings, Donald Struan Robertson (who later became Regius Professor of Greek in the University of Cambridge), Janet Robertson, who later became a portrait painter, and Margaret Robertson (married name Hills), who was a notable suffragist and local politician. Her father gave her regular drawing lessons during her early childhood, which later provided her with the necessary skills to illustrate her scientific publications herself. At the age of eight Robertson began attending the North London Collegiate School founded and run by Frances Buss, one of the leading proponents for girls' education. Under the direction of the school's science teacher Edith Aitken, Robertson discovered a fascination with botany, publishing her first piece of research in 1894 in the school's magazine and later coming first in the school's botany examinations, winning a scholarship. It was here that Robertson first met Ethel Sargant, a plant morphologist who gave regular presentations to the school science club. Sargant would later become her mentor and colleague, having a profound influence on Arber's research interests and methods. In 1897, Robertson began studying at University College, London, gaining her BSc in 1899. After gaining an entrance scholarship Arber became a member of Newnham College, Cambridge and took a further degree in Natural Sciences. She gained first class results in every examination at both universities, along with several prizes and medals from University College, London. After finishing her Cambridge degree in 1902 Robertson worked in the private laboratory of Ethel Sargant for a year, before returning to University College, London as holder of the Quain Studentship in Biology. She was awarded a Doctorate of Science in 1905. Robertson met Edward Alexander Newell Arber (1870–1918) while studying at Newnham College. They married on 5 August 1909, and moved back to Cambridge, where she would remain for the rest of her life. Her only child, Muriel Agnes Arber, was born in 1913, became a geologist, and died in 2004. Arber and her husband had many interests in common, and her marriage was described as 'happy'. Arber was awarded a Research Fellowship from Newnham College in 1912 and published her first book Herbals, their origin and evolution in the same year. Her husband Newall Arber died in 1918 following a period of ill health. Arber never remarried, but continued with her research. She studied in the Balfour Biological Laboratory for Women from her marriage until the laboratory's closure in 1927. Arber maintained a small laboratory in a back room of her house from then until she stopped performing bench research in the 1940s and turned to philosophical study. Agnes Arber died on 22 March 1960, at the age of 81, and is buried in the churchyard of St Andrew's, Girton.

Before attending University College, London Arber spent the summer of 1897 working with Ethel Sargant in her private laboratory in Reigate, where Sargant instructed her on microtechniques used to prepare plant specimens for microscopic examination. Arber returned to work in Sargant's laboratory at least once during the summer holidays while she was studying at University College London. Sargant employed Arber between 1902 and 1903 as a research assistant working on seedling structures, during which time in 1903 she published her first paper 'Notes on the anatomy of Macrozamia heteromera in Proceedings of the Cambridge Philosophical Society. Whilst at University College London Arber conducted research on the gymnosperm group of plants, producing several papers on their morphology and anatomy. The study and philosophy of plant morphology would become the central focus of her later work.

In 1909 Arber was granted space in the Balfour Laboratory for Women by Newnham College. This building had been purchased and founded by the two women's colleges of the university in 1884 for the use of their students and researchers (women at this time were not permitted to attend laboratory demonstrations and practical classes). Arber worked in the laboratory until its closure in 1927. • Illustration — Photo of Arber in 1911 Following the award of a Research Fellowship by Newnham College between 1912 and 1913 Arber published her first book in 1912. Herbals, their origin and evolution describes the transformation of printed Herbals between 1470 and 1670. Arber links the emergence and development of botany as a discipline within natural history with the evolution of plant descriptions, classifications and identifications seen in Herbals during this period. Arber was able to consult the large collection of printed Herbals in the library of the Botany School at Cambridge as part of her research for this work. It was largely re-written and expanded for a second edition published in 1938, was published as a third edition in 1986 and is still considered the standard work for the history of Herbals. • Illustration — Illustration of Limnanthemum from Water Plants A Study of Aquatic Angiosperms by Agnes Arber Arber focused her research on the anatomy and morphology of the monocot group of plants, which she had originally been introduced to by Ethel Sargant. By 1920 she had authored two books and 94 other publications. Her second book Water Plants: A Study of Aquatic Angiosperms was published in 1920. In this book Arber presents a comparative study of aquatic plants by analysing differences in their morphology. Arber also provides interpretations of the general principles she used to create her analysis. Her study was the first to provide a general description and interpretation of aquatic plants. She was an excellent draughtsperson and often illustrated her books herself; she drew about a third of the illustrations in Water Plants. In 1921, the "botanical establishment" denied Arber the 1921 presidency of British association for the advancement of science (BAAS)'s Botany section (Section K). This process was initiated by Frederick Orpen Bower, based on the premise that Arber should not follow Saunders immediately as president as both were women from Cambridge and that there were more senior male botanists up for the presidency. This reflected an antipathy against Arber not only as a female botanist but also as a Cambridge botanist. He claimed that Edinburgh (where the meeting was held) "has the right to expect better than this... To ask Balfour [of Edinburgh] to sit under the Presidency of Mrs. Arber is ridiculous!" Albert Seward met with Arber, who withdrew her candidacy and resigned as the secretary of the BAAS. In 1925 Arber published her third book The Monocotyledons. The Editors of the Cambridge Botanical Handbooks series had asked Ethel Sargant in 1910 to prepare a volume on the monocots for this series. However ill-health and advancing years made it almost impossible for Sargant to complete the book, and in 1918 she suggested Arber to complete the work. The Monocotyledons continues Arber's morphological methods of analysis she presented in Water Plants. She provides a detailed study of the monocot plants from comparing their internal and external anatomy. However her discussion of the general principles she uses in her analysis are more explicit in this volume, as she discusses the methods and philosophy of morphological study. Although comparative anatomical analysis as demonstrated in The Monocotyledons and Water Plants: A Study of Aquatic Angiosperms was central to botanical investigation in the early 20th century, there were distinct differences between British and European researchers concerning the aims of morphological study. Arber addressed this by creating a distinction between "pure" and "applied" morphology, with her work focusing on comparative anatomy to investigate questions concerning significant topics such as constructing phylogenies, instead of using traditional views of plant structure. This view was further developed in her later work.

After the 1927 closure of the Balfour Laboratory Arber set up a small laboratory in a back room of her house to conduct her research, after the resident head of the Botany School Professor Albert Seward claimed there was no space in the School for Arber to continue her research using its facilities. Arber had been introduced to the idea of private research from her time spent with Ethel Sargant in 1902–1903, and from later comments to members of Girton College Natural Sciences club and in letters to friends she stated she liked working at home due to challenges posed by independent research, despite not originally making the choice herself. After the publication of The Monocotyledons Arber continued her research into this group, concentrating her research into the Gramineae family of plants, especially cereals, grasses and bamboo. This led to the publication of her final book concerning plant morphology, The Gramineae in 1934. In this book Arber described the life cycles, embryology and reproductive and vegetative cycles of cereals, grasses and bamboo using comparative anatomical analysis of these plants. Recognising the importance of these plants to the development of human societies, Arber begins this study with the history of these plants in relation to humans, with "the more strictly botanical aspect is treated as developing out of the humanistic". The book was preceded by 10 papers in The Annals of Botany detailing the results of her research. Between 1930 and 1942 Arber conducted research into the structure of flowers, where she investigated the structure of many different forms and used morphological information to interpret other flower structures. Her results were published in 10 review papers spanning this period. In 1937 she published a summary of the morphological ideas which had been discussed concerning floral structure, which was considered an important review article for morphological studies. In January 1942 Arber published her last paper involving original botanical research. All of her subsequent publications were entirely concerned with historical and philosophical topics.

Agnes Arber was recorded at birth as Agnes Robertson. The authorities additionally record the headings A. Arber, Agnes Robertson Arber and A. Her recorded languages were German and English. She was the child of Henry Robert Robertson. She married Edward Alexander Newell Arber.

Agnes Arber worked in botany, plant morphology, plant anatomy, history of botany and bibliography. She was employed by University College London and Balfour Biological Laboratory. She belonged to Royal Society, Linnean Society of London, Botanical Society of America, British Science Association and International Society of Plant Morphologists.

Works recorded as notable number 3: Herbals, their origin and evolutiion; a chapter in the history of botany, 1470-1670, Water plants; a study of aquatic angiosperms and The Natural Philosophy of Plant Form.

Goethe's botany: the Metamorphosis of plants (1790) and Tabler's Ode to nature (1782) / with an introduction and translations by Agnes Arber. Herbals, their origin and evolution: a chapter in the history of botany, 1470-1670 / by Agnes Arber (Mrs. The Agnes Arber collection / James D. Agnes Arber, 1879-1960 / [H. Agnes Arber is recorded as botanist, translator, geneticist and historian.